1981 Biology Sciences, Universidad Complutense de Madrid (Spain)

1982 Master of Sciences, Universidad Complutense de Madrid (Spain).

1985 Ph.D. in Biochemistry, Universidad Complutense de Madrid (Spain).

1987 Postdoctoral, Center of Mollecuar Biology (CSIC-UAM), Universidad Autónoma de Madrid (Spain).

1991 Postdoctoral, Cambridge University (Great Britain).

2001 present Head of Dpt., Dpt. Structural & Computational Biology, EMBL (Germany).

2005 present ICREA Professor and Group Leader at the Center for Genomic Regulation, Barcelona (Spain)